Principal Research Fellow and Honorary Consultant Neurologist at University College London. Originally from Renfrewshire in Scotland he trained in Glasgow, London, Brussels and San Francisco. He now works within the Dementia Research Centre at the UCL Queen Square Institute of Neurology 70% of the time, and spends the other 30% of his week doing clinical work, he has close collaborations with groups at Washington University, St Louis and the University of Gothenburg - all in his capacity as a clinical (cognitive) neurologist and biomarker researcher with a particular interest in measuring protein turnover in humans.